Daphne seeks to establish her career and auditions for a country music singing competition. Scarlett invites Sean to see live music. Avery makes a decision. The band gets invited to perform on late night TV. Will’s choice catches up with him.
Daphne and Jake perform at school; Deacon and Jessie run into Brad; Alannah’s stage presence grows, creating a division in the band; Gunnar questions his relationship with Alannah; Maddie joins Jonah in Miami and bonds with Twig; Avery visits Juliette.
Daphne struggles to accept Deacon and Jessie’s relationship; the new addition to the boys’ band becomes a hit; Gunnar falls back into old habits; Scarlett takes an interest in a veteran undergoing equine therapy.
Darius helps Juliette unlock a dark truth about her past; Daphne and Jake bond over each other’s dislike of their parents dating; Scarlett volunteers at an equine therapy ranch; Gunnar, Will and Avery struggle to find their sound.
With Deacon and the artists of Highway 65 taking a stand, Zach sends a message. Maddie and Juliette’s conflict comes to a head. Daphne goes to a pool party. Gunnar returns from Avery’s tour.
Juliette shoots a mascara commercial but takes issue with the direction; Deacon gets invited to play the Opry, but hesitates; Avery and Gunnar embrace life on the road.